Edinburgh Zoo's panda Yang Guang has had both testicles removed after the discovery of tumours.

The panda, also known as 'Sunshine', underwent surgery earlier this month after group consultation by British and Chinese giant panda specialists.

Staff confirmed the news on Tuesday morning but said the tumours have not been a factor in Yang Guang and female panda Tian Tian's struggles to have a cub.

Darren McGarry, head of Living Collections at Edinburgh Zoo, said, "We are pleased Yang Guang is recovering well.

"Testicular tumours have previously been reported in giant pandas.

"The tumours we removed developed recently, so they have not been a factor in Yang Guang and Tian Tian not having had a cub.

"We will discuss next steps, including future breeding possibilities, with our partners in China over coming weeks."

Unsuccessful efforts have been made to breed Tian Tian and Yang Guang since the pair arrived in 2011 as part of a conservation breeding programme.

In September, 2017, Tian Tian failed to produce a cub after a fifth attempt.